speciesaccounts_whooperswanWhooper Swan

Cygnus cygnus

The Iceland population of Whooper Swan breeds exclusively in Iceland and winters primarily in Britain and Ireland, with smaller numbers remaining in Iceland and some also migrating as far as the near European continent.

There are four other populations of Whooper Swan: i) the Northwest European population breeds in Scandinavia and north European Russia and winters in northwest and central Mainland Europe; ii) the Black Sea and east Mediterranean population breeds in northern Europe and western Siberia and winters in the Black Sea and east Mediterranean; iii) the Caspian and central Asia population breeds in west and central Siberia and winters in the Caspian and central Asia; and iv) the east Asia population breeds in central and east Siberia to northeast China and winters in east Asia.

Whooper Swans undertake what is probably the longest sea crossing of any swan species, migrating 800-1,400 km between Britain/Ireland and Iceland.

The Goose & Swan Monitoring Programme only monitors the Iceland population of Whooper Swan.

  • Conservation Status

    Global status (IUCN Red List of Threatened Species) Least Concern
    African-Eurasian Waterbird Agreement (AEWA) B1
    European status (European Red List of Birds) Least Concern (Europe and EU27)*
    The Birds Directive (European Commission) Annex I
    UK status (Birds of Conservation Concern) Amber
    UK quarry species (Wildlife & Countryside Act 1981) not huntable

    Population Status

    Flyway population size (CSR 7; Wetlands International 2015) 34,000 individuals
    UK estimate (APEP 4) 19,500 individuals
    GB estimate (Frost et al. 2019) 16,100 individuals
    All-Ireland estimate (Burke et al. 2018) 15,370 individuals
    UK trend (Frost et al. 2020) 25-year trend (1992/93-2017/18) = 210% increase
    10-year trend (2007/08-2017/18) = 43% increase

    Summary statistics

    Table 1. Census-derived estimates of the population size of Iceland Whooper Swan, 1986-2015; recorded during the International Swan Census.

    Census year Estimate of population size
    2015 34,004
    2010 29,3001
    2005 26,366
    2000 20,856
    1995 15,842
    1991 18,035
    1986 16,742

    1Since the publication of January 2010 census results (Hall et al. 2012), additional counts totalling 68 birds were received from four sites in the Republic of Ireland resulting in a revised 2010 population estimate of 29,300 (previously 29,232).

    Table 2. Annual estimates of the percentage of young (%) and mean brood size (young per successful pair) of Whooper Swan in Britain (recorded at WWT Caerlaverlock, WWT Martin Mere & Ribble Estuary, and the Ouse Washes, where data are collected annually), 2003/04-2019/20.

    Season Percentage of young (%) Mean brood size
    2019/20 13.6 2.0
    2018/19 14.7 1.8
    2017/18 17.9 2.0
    2016/17 11.7 2.0
    2015/16 11.8 1.9
    2014/15 16.0 2.0
    2013/14 13.3 1.9
    2012/13 13.0 1.9
    2011/12 10.6 2.1
    2010/11 14.4 1.9
    2009/10 13.0 2.1
    2008/09 14.9 2.6
    2007/08 20.4 2.5
    2006/07 13.8 2.4
    2005/06 10.6 2.4
    2004/05 13.7 2.4
    2003/04 15.0 2.7
    References

    Burke, B., L.J. Lewis, N. Fitzgerald, T. Frost, G.E. Austin & T.D. Tierney. 2018. Estimates of waterbird numbers wintering in Ireland, 20011/12-2015/16. Irish Birds 11: 1-12

    Frost, T.M., N.A. Calbrade, G.A. Birtles,  H.J. Mellan, C. Hall, A.E. Robinson,S.R. Wotton, D.E. Balmer & G.E. Austin. 2020. Waterbirds in the UK 2018/19: The Wetland Bird Survey. BTO, RSPB and JNCC, in association with WWT. British Trust for Ornithology, Thetford.

    Frost, T., G.E. Austin, R.D. Hearn, S. McAvoy, A. Robinson, D.A. Stroud, I. Woodward & S.R. Wotton. 2019. Population estimates of wintering waterbirds in Great Britain. British Birds 112: 130-145.

    Data access

    Whooper Swan data presented in in Table 1 (Estimate of population size) and Table 2 (Percentage of young / Mean brood size in Britain) are licensed under the Open Government Licence 3.0 except where otherwise stated.

    When you use information from this report under the Open Government Licence you must include the following attribution:

    Contains Goose & Swan Monitoring Programme (GSMP) data from “WWT. 2020. Goose & Swan Monitoring Programme: survey results for Whooper Swan Cygnus cygnus. WWT/JNCC/NatureScot, Slimbridge” retrieved from https://monitoring.wwt.org.uk/our-work/goose-swan-monitoring-programme/species-accounts/whooper-swan/ © copyright and database right 2020. The GSMP is organised by the Wildfowl & Wetlands Trust (WWT) in partnership with the Joint Nature Conservation Committee (JNCC) and NatureScot with fieldwork conducted by volunteers.

  • Iceland Whooper Swans breed throughout Iceland (Garðarsson & Skarphedinsson 1984). Large flocks of non-breeders and failed breeders gather on marine and brackish waters in Iceland during late summer, and the majority leave Iceland from mid October to mid November; ringing and count programmes indicate that most migrate to winter in Britain and Ireland, with a small proportion remaining to overwinter in Iceland (Black & Rees 1984, Gardarsson 1991, Rees et al. 2002). Sites in northern Britain and Ireland act as major landfalls, e.g. Shetland, Orkney and, especially, Loughs Swilly and Foyle in Northern Ireland (Robinson et al. 2004).

    flyway map whooper

    Flyway of Iceland Whooper Swan

    Spring migration back to Iceland starts in March and April. Satellite tacking studies have shown that the swans may fly either direct from Ireland to Iceland, a 1,400 km flight overseas, or via Scotland (Rees 2009), though there is a minimum 800 km flight between Britain and Iceland and the birds are susceptible to strong head or side winds blowing them off course (Pennycuick et al. 1996, Rees 2009).

    In Ireland, Whooper Swans are distributed throughout, with highest numbers usually found at Loughs Neagh & Beg (Antrim, Armagh, Down, Londonderry, Tyrone), Lough Foyle (Donegal, Londonderry) and Upper Lough Erne (Fermanagh).

    Two key sites in England regularly hold the majority of wintering Whooper Swans in Britain: WWT Martin Mere & surrounding area (Lancashire) and Ouse Washes (Cambridgeshire, Norfolk). In Scotland, birds are widely distributed; from Orkney down the north and western coast, across to the Outer Herbrides, and down through eastern and central counties to the Solway Estuary in the southwest – this latter site usually holding the highest numbers. Swans are sparsely distributed elsewhere in England and Wales.

    Although the Iceland population is protected throughout its range, illegal shooting remains a threat, with 13% of live Whooper Swans x-rayed since 2000 found with shotgun pellets embedded in their body tissue (Newth et al. 2011). Shooting has also accounted for the death of 8.3% of dead Whooper Swans recovered along their migration route since the 1980s (Newth et al. 2011). Illegal shooting is known to have occurred in all countries within the species range, including the UK (Rees et al. 2002, Newth et al. 2011).

    There is evidence of some interchanges between the Iceland and Northwest European populations. Ring sightings of British ringed birds in Norway, Denmark and the Netherlands suggest that up to 600 birds from the Iceland population may winter in mainland Europe while a small number of Finnish ringed birds have been sighted in Britain giving rise to the suggestion that around 200 birds from the Northwest Europe population visit Britain (Garðarsson 1991, Cranswick et al. 1997, Laubek et al. 1998).

    References

    Cranswick, P.A., J.M. Bowler, S. Delany, Ó. Einarsson, A. Garðarsson, J.G. McElwaine, E.C. Rees & J.H. Wells. 1997. Whooper Swans wintering in Britain, Ireland and Iceland: International Census, January 1995. Wildfowl 47: 17-30.

    Garðarsson, A. 1991. Movements of Whooper Swans Cygnus cygnus neckbanded in Iceland. In: Sears, J. & P.J. Bacon (eds.). Proceedings of the 3rd IWRB International Swan Symposium, Oxford, 1989. Wildfowl Special Supplement No. 1: 189-194.

    Gardarsson, A. & K.H. Skarphedinsson. 1984. A census of the Icelandic Whooper Swan population. Wildfowl 35: 37–47.

    Laubek, B., H.L. Knudsen & A. Ohtonen. 1998. Migration and winter range of Whooper Swans Cygnus cygnus breeding in different regions of Finland. In: Laubek, B. 1998. The Northwest European Whooper Swan (Cygnus cygnus) population: ecological and management aspects of an expanding waterfowl population. PhD thesis, University of Aarhus, Denmark.

    Newth, J.L., M.J. Brown & E.C. Rees. 2011. Incidence of embedded shotgun pellets in Bewick’s swans Cygnus columbianus bewickii and whooper swans Cygnus cygnus wintering in the UK. Biological Conservation 144: 1630-1637.

    Pennycuick, C.J., Ó. Einarsson, T.A.M. Bradbury & M. Owen. 1996. Migrating Whooper Swans Cygnus cygnus: satellite tracks and flight performance calculations. Journal of Avian Biology 27: 118–134.

    Rees, E.C. 2009. Lough Neagh Wetlands Whooper Swan Study: Project Report. WWT Report to Irish Whooper Swan Study Group, Wildfowl & Wetlands Trust, Slimbridge, UK.

    Rees, E.C., K. Colhoun, Ó. Einarsson, J.G. McElwaine, A. Petersen & S. Thorstensen. 2002. Whooper Swan Cygnus cygnus. In: Wernham, C.V., M.P. Toms, J.H. Marchant, J.A. Clark, G.M. Siriwardena & S.R. Baillie (eds.) The Migration Atlas: Movements of the Birds of Britain and Ireland. pp. 154–157. T. & A.D. Poyser, London, UK.

  • International Swan Census

    A coordinated, international census of the Iceland Whooper Swan population is undertaken in Britain, Ireland and Iceland every five years. The census covers many non-wetland and/or temporarily flooded areas that are not routinely covered by other annual schemes.

    Results from the censuses are presented in various reports and papers. See our Reports & newsletter page.

    Find out more about the International Swan Census

    Wetland Bird Survey and Irish Wetland Bird Survey

    Whooper Swan numbers in the UK and Republic of Ireland are monitored annually through the Wetland Bird Survey (WeBS) and the Irish Wetland Bird Survey (I-WeBS), respectively. Although these schemes provide good coverage of a high proportion of wintering sites used by Whooper Swans, a number of non-wetland or temporally flooded areas are not surveyed, hence a more comprehensive census, that focuses specifically on Whooper Swans, is required to fully access the size of the population; such a survey is undertaken every five years (see above).

    GSMP age assessments

    The annual breeding success of the Iceland population of Whooper Swan is monitored in Britain through age assessments that are undertaken throughout the autumn and early winter. Counters record the number of first winter birds present within a flock and individual brood sizes (i.e. how many young in each family group).

    Results from these age assessments are presented on the ‘Latest results’ tab, and a summary table can also be found on the ‘Status summary’ tab.

    Find out more about age assessments

  • Results for 2019/20 [added October 2020]

    Abundance

    WeBS/I-WeBS

    The abundance of Whooper Swan in the UK and the Republic of Ireland in 2019/20 was monitored through the Wetland Bird Survey (WeBS) and the Irish Wetland Bird Survey (I-WeBS), respectively. Results from these schemes are presented in reports which are available via the schemes’ websites.

    International Swan Census

    The International census of the Icelandic Whooper Swan population is carried out every five years. The 8th census was carried out in January 2020 and the results will be available in autumn 2021.

    The census is organised overall by the IUCN SSC Swan Specialist Group, and coordinated in Britain, Ireland and Iceland by WWT in partnership with Birdwatch Ireland, the Irish Whooper Swan Study Group and colleagues in Iceland.

    The 7th international census took place in January 2015. A total of 34,004 Whooper Swans was recorded, representing an increase of 16% since the previous census in 2010 (Figure 1). The results from this census have been presented in Hall et al. (2016).

    Figure 1: The number of Icelandic Whooper Swans recorded during the International Swan Census, 1986–2015. Note: Wales and the Isle of Man are combined as each holds less than 1% of the total population.

    Breeding success

    Whooper Swan age assessments were conducted in fifteen regions across Britain and Ireland during winter 2019/20 (Table 1). Age assessments were made in all regions in mid-winter (9–15 January 2020), to coincide with the International Swan Census when more flocks were covered across the flyway and when the majority of families were likely to have arrived from Iceland to wintering sites (Rees et al. 1997). Regional variation in the percentage of young and mean brood size was assessed to determine any bias in the geographical distribution of family parties.

    A total of 30,185 Whooper Swans was aged (88.8% of the total population; Hall et al. 2016): 12,423 birds in England, 3,285 in Scotland, 193 in Wales, 27 in the Isle of Man, 4,125 in Northern Ireland and 10,132 in the Republic of Ireland (Table 1). Overall, 17.1% of birds were cygnets, this being higher than the previous ten-year mean for Whooper Swans wintering at sites in Britain and Ireland (15.8% ± 0.6 SE for 2009/10–2018/19). The mean brood size for pairs with young was 2.03 which was slightly higher than the previous ten-year mean (2.00 ± 0.03 SE for 2009/10–2018/19).

    Table 1: The percentage of young (%) and mean brood size of Whooper Swans during the 2019/20 winter (regions defined below).

    Region Total aged (number of young) Percentage of young (%) Number of broods (number of young) Mean brood size
    North & Northeast England 242 (35) 14.5 14 (33) 2.36
    Northwest England 2,478 (494) 19.9 149 (387) 2.6
    Central England 434 (93) 21.4 3 (3) 1
    East Central England 9,264 (1,133) 12.2 582 (1,078) 1.85
    Southwest England 5 (0) 0
    North Scotland 1,474 (336) 22.8 57 (138) 2.42
    Central Scotland 497 (103) 20.7 45 (89) 1.98
    East Scotland 157 (23) 14.6 7 (18) 2.57
    West Scotland 182 (38) 20.9 16 (33) 2.06
    South Scotland 975 (154) 15.8 56 (119) 2.13
    Isle of Man 27 (5) 18.5 3 (5) 1.67
    North Wales 125 (20) 16 6 (14) 2.33
    Northwest Wales 68 (19) 27.9
    Northern Ireland 4,125 (825) 20 373 (749) 2.01
    Republic of Ireland 10,132 (1,888) 18.6 715 (1,438) 2.01
    Overall 30,185 (5,166) 17.1 2,206 (4,104) 2.03

    Regions (counties from which data were received in 2019/20):

    • North & Northeast England: Yorkshire, Northumberland
    • Northwest England:  Lancashire (WWT Martin Mere/Ribble Estuary), Cumbria
    • Central England: Cheshire, Shropshire, Nottinghamshire
    • East Central England: Cambridgeshire and Norfolk (WWT Welney/Ouse Washes/Nene Washes), Lincolnshire, Suffolk
    • Southwest England: Gloucestershire
    • Southwest Scotland: Dumfries & Galloway (WWT Caerlaverock)
    • North Scotland: Aberdeenshire, Sutherland, Shetland, Ross & Cromarty, Outer Hebrides, Orkney, Moray, Inverness-shire, Caithness
    • Central Scotland: Stirlingshire, Perthshire, Midlothian, Lanarkshire
    • East Scotland: Fife
    • West Scotland: Argyll and Bute, Renfrewshire, Kintyre, Ebudes, Dunbartonshire, Clyde
    • South Scotland: Wigtownshire, Selkirkshire, Roxburghshire, Kirkudbrightshire, Borders, Ayrshire
    • Isle of Man
    • North Wales: Anglesey, Flintshire, Carmarthenshire, Caernarvonshire
    • Northwest Wales: Merionethshire
    • Northern Ireland: Co. Antrim, Co. Armagh, Co. Down, Co. Fermanagh, Co. Londonderry, Co. Tyrone
    • Republic of Ireland: Co. Cavan, Co. Clare, Co. Cork, Co. Donegal, Co. Galway, Co. Kerry, Co. Leitrim, Co. Limerick, Co. Mayo, Co. Meath, Co. Monaghan, Co. Offaly, Co. Roscommon, Co. Sligo, Co. Tipperary, Co. Waterford, Co. Wexford and Co. Wicklow

    There was evidence of variation in the distribution of families between regions (X212 = 264.8, P < 0.05). Overall, higher breeding success was found for those wintering in more northerly regions (Scotland and Northern Ireland) compared to those in more southerly regions (England, IoM, Wales and the Republic of Ireland) (24.9 %, n = 7,410 and 19.3 %, n = 22,775, respectively) (X21 = 55.8, P = < 0.05).

    Age assessments of Whooper Swans have been regularly undertaken at and around WWT centres (WWT Welney/Ouse and Nene Washes, WWT Martin Mere/Ribble Estuary and WWT Caerlaverock) since the early 1980s. In 2019/20, the mean percentage of young in flocks at these sites combined, was 13.6% (11,432 birds aged) (Figure 2), which is the same as the previous ten-year mean (2009/10–2018/19; 13.6 % ± 0.69 SE); whilst the mean brood size was 2.00 cygnets per family, this being higher than the previous ten-year mean (2009/10–2018/19; 1.96 ± 0.03 SE).

    At the individual sites, the percentage of young was highest amongst flocks at WWT Martin Mere/Ribble Estuary (19.9%), with 17.6% and 12% recorded at WWT Caerlaverock and WWT Welney/Ouse and Nene Washes, respectively (Figure 3). Mean brood size was, however, highest at WWT Caerlaverock (2.92 cygnets per family), whilst 2.58 cygnets and 1.84 cygnets were recorded at WWT Martin Mere/Ribble Estuary and WWT Welney/Ouse and Nene Washes, respectively.

    Figure 2: The percentage (%) of young (blue circles), with the rolling five-year mean (red line), and mean brood size (green triangles) of Whooper Swans recorded at WWT Welney/Ouse and Nene Washes, WWT Caerlaverock and WWT Martin Mere/Ribble Estuary, 2005/06–2019/20. Five-year mean values for the percentage of young were calculated for the five years preceding the year in question (e.g. mean presented for 2019/20 is for 2014/15–2018/19). 

    Figure 3. The percentage of young Whooper Swans recorded at WWT Martin Mere/Ribble Estuary, WWT Caerlaverock, WWT Welney/Ouse Washes and Nene Washes (Britain), Northern Ireland and the Republic of Ireland, 2005/06–2019/20.  

    Discussion

    In 2019, Icelandic Whooper Swans experienced a fairly average breeding season. The overall percentage of young for flocks wintering in Britain and Ireland was only slightly higher (by c.1%) than the previous ten-year mean; while the percentage of young recorded amongst flocks at and around WWT centres (for which there is a long-term dataset) was the same as the previous ten-year mean.

    The mean temperature (9.3˚C) for Akureryi, a key breeding area, in June 2019, was slightly lower than the June temperature recorded in the previous five years (2014-2018; 10.2 ˚C; Tutiempo 2020); however, no extreme conditions were reported from the breeding grounds and, therefore, unlikely to have impacted the swans’ breeding success.

    Higher breeding success was found in northern regions of Britain and Ireland compared with southern regions which may reflect a general preference for Whooper Swan families to select wintering sites closest to their Icelandic breeding grounds (Rees et al. 1997).

    Acknowledgements

    Special thanks to all observers and Local Organisers who took part in the international census and the productivity surveys, including the WeBS, I-WeBS, Irish Whooper Swan Study Group and GSMP networks. We are especially grateful to Brian Burke and Graham McElwaine for coordinating the surveys across Ireland.

    References

    Hall, C., O. Crowe., G. McElwaine., O. Einarsson., N. Calbrade & E. Rees. 2016. Population size and breeding success of the Icelandic Whooper Swan Cygnus cygnus: results of the 2015 international census. Wildfowl 66: 75–97.

    Rees, E.C., J.S. Kirby & A. Gilburn. 1997. Site selection by swans wintering in Britain; the importance of habitat and geographic location. Ibis 139: 337–352.

    Tutiempo. 2020. https://en.tutiempo.net/climate/06-2019/ws-40630.html. [Accessed July 2020.]

    Data access

    Whooper Swan abundance data for Britain presented in this report are licensed under the Open Government Licence 3.0 except where otherwise stated.

    Whooper Swan breeding success data for Britain presented in this report, with the exception of data for 2018/19 and 2019/20 presented in Figure 3 (percentage of young/mean brood size for Bewick’s Swans recorded at WWT centres), are licensed under the Open Government Licence 3.0 except where otherwise stated.

    When you use information from this report under the Open Government Licence you must include the following attribution:

    Contains Goose & Swan Monitoring Programme (GSMP) data from “WWT. 2020. Goose & Swan Monitoring Programme: 2019/20 survey results for Whooper Swan Cygnus cygnus. WWT/JNCC/NatureScot, Slimbridge” retrieved from  https://monitoring.wwt.org.uk/our-work/goose-swan-monitoring-programme/species-accounts/whooper-swan/ © copyright and database right 2020. The GSMP is organised by the Wildfowl & Wetlands Trust (WWT) in partnership with the Joint Nature Conservation Committee (JNCC) and NatureScot with fieldwork conducted by volunteers.

  • Previous annual results will be archived here. Published results from the International Swan Census can be found on the Reports & newsletter page.

    2018/19 Results

    2017/18 Results

    2016/17 Results

    2015/16 Results 

    2014/15 Results

    2013/14 Results

    2012/13 Results

    2011/12 Results

    2010/11 Results

    2009/10 Results

    2008/09 Results

    2007/08 Results

    2006/07 Results

    2005/06 Results

  • Background

    Regular colour-marking of Whooper Swans in Britain started in 1980, following the construction of swan-pipes at WWT Caerlaverock and WWT Welney. A swan-pipe is a long, netted trap which is baited with food to encourage birds to enter. A gate is dropped to close off the entrance and the birds are moved into a holding area, before being caught, ringed and released.

    Catches at WWT Caerlaverock have resulted in X Whooper Swans being ringed, and a third swan-pipe at WWT Martin Mere, constructed in 1990, has also led to the capture and ringing of several hundred birds. During the 1980s, the number of Whooper Swans wintering at WWT Welney was relatively low (100-300 birds), and by the winter of 1994/95, when over a thousand birds were first recorded at the site, the swan-pipe had fallen into disrepair, with fewer than 50 swans having been caught (Rees et al. 2002). In 2007, a new swan-pipe was built at the centre and since then over 200 birds have been captured and ringed.

    Whooper Swans have also been caught using cannon-nets in the fields around WWT Welney, although this tends to result in fewer birds being captured but has resulted in x birds ringed.

    Smaller numbers of Whooper Swans have been colour-ringed elsewhere in Britain, including South Uist in the late 1980s, northeast England throughout the 1990s, southwest Wales in 1991 and in Northern Ireland, particularly at Castle Espie, since winter 1992/93 (Rees et al. 2002) and more recently small numbers have been caught by cannon netting in north Wales and north Yorkshire.

    In Iceland, the Whooper Swan ringing programme, using metal rings, started in the 1960s. In the early- to mid- 1980s, a few hundred birds from moulting flocks were marked with neck-collars by Arnthor Gardarsson (Rees et al. 2002). In 1988, WWT in collaboration with Icelandic colleagues caught and colour-ringed Whooper Swans on their breeding territories (Rees & Bowler 1996), and since then annual trips have been made to Skagafjörður, Mývatnsheiði and Jökuldalsheiði in northern Iceland to ring breeding and non-breeding birds. Since 2009, this work has solely been undertaken by Ólafur Einarsson and Sverrir Thorstensen in conjunction with the Icelandic Institute of Natural History.

    Resightings of colour-marked Whooper Swans have lead to a number of important discoveries including:

    • Confirmation that swans move frequently between Britain and Ireland during the winter. Northern parts of Ireland, in particular, are used as staging sites by birds wintering in Britain, as well as those wintering elsewhere in Ireland (McElwaine et al. 1995).
    • Evidence that birds breeding in the more western parts of Iceland (in Skagafjörður) concentrate largely in more western parts of the wintering range (Ireland), whereas those ringed in east Iceland (Norður-Múlasýsla) were more likely to occur in Scotland and in continental Europe (Gardarsson 1991, McElwaine et al. 1995, Newth et al. 2007).
    • Indications of an overall southerly shift in distribution as the winter progressed (McElwaine et al. 1995, Newth et al. 2007).

    WWT’s colour-ringing

    Whooper Swans are caught and ringed at WWT Caerlaverock (Dumfries & Galloway), WWT Martin Mere (Lancashire) and WWT Welney (Norfolk). This is undertaken as part of WWT’s long-term life-history study on the species. Each bird is fitted with a coloured leg ring, as well as a metal ring and various body size measurements and samples are also taken to assess its condition and health. By catching these birds we have been able to identify issues affecting them, such as illegal shooting and lead poisoning (Newth et al. 2011, Newth et al. 2012). This information helps to inform subsequent conservation measures taken to reduce the scale of these threats.

     Ring combinations used
    Yellow leg rings (black lettering) Red leg rings (white lettering) Orange leg rings (black lettering)
    Alpha/alpha/alpha AJA – VZZ AAA – BLZ ZAA – ZZZ
    Numeric/alpha/alpha 3AA – 9ZZ
    Alpha/alpha/numeric FA3 – ZZ9
    Numeric/numeric/alpha 31A – 99Z
    Alpha/numeric/numeric A33 – Z99
    Alpha/numeric/alpha A3A – Y9Z

    If you see any colour-marked Whooper Swans with these combinations please send your sightings to colourmarkedwidlfowl@wwt.org.uk

    Project partners

    We work with the Icelandic Institute of Natural History, the Icelandic Research Council, the Irish Whooper Swan Study Group, Olafur Einarsson, Sverrir Thorstensen, the University College Cork and the University of Iceland.

    References

    Garðarsson, A. 1991. Movements of Whooper Swans Cygnus cygnus neckbanded in Iceland. In: Sears, J. & P.J. Bacon (eds.). Proceedings of the 3rd IWRB International Swan Symposium, Oxford, 1989. Wildfowl Special Supplement No. 1: 189-194.

    McElwaine, J.G., J.H. Wells & J.M. Bowler. 1995. Winter movements of Whooper Swans visiting Ireland: preliminary results. Irish Birds 5: 265-78.

    Mitchell, C. & M. Ogilvie. 1997. Fifty years of wildfowl ringing by the Wildfowl & Wetlands Trust. Wildfowl 47: 240-247.

    Newth, J., K. Colhoun, Ó. Einarsson, R. Hesketh, J.G. McElwaine, S. Thorstensen, A. Petersen, J. Wells & E.C. Rees. 2007. Winter distribution of Whooper Swans (Cygnus cygnus) ringed in four geographically discrete regions in Iceland between 1988 and 2006: an update. Wildfowl 57: 98-119. Download.

    Rees, E.C. & J.M. Bowler. 1996. Fifty years of swan research and conservation by the Wildfowl & Wetlands Trust. Wildfowl 47: 248-263. Download.

    Rees, E.C., K. Colhoun,  Ó. Einarsson, J.G. McElwaine, Æ. Petersen & S. Thorstensen. 2002. Whooper Swan Cygnus cygnus. In: Wernham, C.V., M.P. Toms, J.H. Marchant, J.A. Clark, G.M. Siriwardena & S.R. Baillie (eds.) The Migration Atlas: Movements of the Birds of Britain and Ireland. Pp. 154-157. T. & AD Poyser, London.

  • Survey results

    International Swan Census reports: See the Reports & newsletter page

    Wetland Bird Survey Alerts

    Wetland Bird Survey annual report

    Relevant publications

    Griffin, L., E.C. Rees & B. Hughes. 2010. The Migration of Whooper Swans in Relation to Offshore Wind Farms. WWT Final Report to COWRIE Ltd, WWT, Slimbridge.

    Kear, J. 2005. Ducks, geese & swans. Oxford University Press.

    Robinson, J.A., K. Colhoun, J.G. McElwaine & E.C. Rees. 2004. Whooper Swan Cygnus cygnus (Iceland population) in Britain and Ireland 1960/61 – 1999/2000. Waterbird Review Series, The Wildfowl & Wetlands Trust/Joint Nature Conservation Committee, Slimbridge. Download

    Other relevant material

    BirdLife International Species factsheet

    British Trust for Ornithology: Bird Facts